Around 1900, fishermen in the Chesapeake Bay area landed more than seventeen million pounds of shad in a single year, but by 1920, over-fishing and the proliferation of milldams and culverts that blocked shad from migrating up their spawning streams had reduced landings to less than four million pounds.
Q:
Which one blocked shad migrations up their spawning streams? Is it ''over-fishing and the proliferation of milldams and culverts'' or ''milldams and culverts'' or ''the proliferation'' only? Thanks